West Africa; Sudanian regions; Ley pasture; Forage; Legumes; Grasses; Afrique de l'ouest; Régions soudaniennes; Jachère fourragère; Fourrage; Légumineuses; Graminées
Abstract :
[en] In Sudanian region of Africa, agricultural systems are under increasing pressure because of human and animal population growth, climate changes, extensive practises, decreasing prices for cash crops, etc. Possibilities of intensification in smallholder farming systems of the Borgou region in Benin are limited due to difficulties to pay for external inputs. Therefore, rural communities rely heavily on low input technologies to increase crop production and animal feed. Cultivated forages are of better-feed quality for ruminants compared to weed fallows. Their integration in cropping system through ley pastures has the potential to increase not only animal feed availability but also to improve soil fertility. This paper reviews some possible grass and legume species that can be used for that purpose with a special focus on the strengths and weaknesses of the species in terms of soil and climate suitability, forage production, nutritive value and soil fertility restoration. The choice of one or several among them as leys in pure stands or mixed forage crops must be taken carefully considering the balance between advantages and disadvantages of the species, the available financial and technical inputs and the adaptation to the local environment. [fr] Dans la région soudanienne de l’Afrique, les systèmes de production agricole sont sous pression
en raison de l’augmentation de la population humaine et animale, des changements climatiques, des modes extensifs de production agricole, de la chute des coûts des produits d’exportation, etc. Les possibilités d’intensification des systèmes de production agricole dans le Borgou au Bénin sont limitées notamment à cause des faibles capacités d’investissement financier des paysans. Par conséquent, les communautés rurales comptent sur des technologies relativement peu coûteuses pour accroître les productions et assurer l’alimentation des animaux. Les cultures fourragères sont des aliments de bonne qualité en comparaison avec les jachères naturelles. Leur intégration dans les systèmes de production sous forme de jachère fourragère offre des possibilités d’augmenter non seulement les disponibilités alimentaires du bétail mais aussi d’accroître la fertilité des sols. Ce document présente une gamme de graminées et de légumineuses susceptibles d’être utilisées en jachère fourragère, mettant l’accent sur les forces et faiblesses de ces espèces en termes d’adaptation au climat, de leur production fourragère, de leur valeur alimentaire et de la restauration de la fertilité des sols. Le choix d’une ou de plusieurs de ces espèces pour les utiliser en mono-culture ou en mélange sous forme de jachère fourragère doit être fait avec précaution en tenant compte des avantages et inconvénients de ces espèces, de la capacité financière des paysans et de leur adaptation à l’environnement local.
Disciplines :
Agriculture & agronomy Animal production & animal husbandry
Author, co-author :
Adjolohoun, Sébastien; Université d'Abomey - Calavi - Faculté des Sciences Agronomiques, Cotonou, Bénin > Département des Productions Animales
Adegbola A.A., 1964. Forage crop research and development in Nigeria. Niger. Agric. J., 1, 34-39.
Adjanohoun E.J. et al., 1989. Contribution aux etudes ethnobotaniques et floristiques en République Populaire du Bénin. Médecine traditionnelle et pharmacopée. Paris: Agence de Coopération Culturelle et Technique (ACCT).
Adjei M.B. & Fianu F.K., 1985. The effect of cutting interval on the yield and nutritive value of some tropical legumes on the coastal grassland of Ghana. Trop. Grassl., 19, 164-171.
Alkoiret T.I., Soule Manne A.A., Gbangboché A.B. & Attakpa E.Y., 2007. Performances d’embouche des ovins Djallonké complémentés avec les coques de graines de coton au Bénin. Livest. Res. Rur. Dev., 19(10), http://www.cipav.org.co/lrrd/lrrd19/10/alko19141.htm, (15/02/08).
Anoka U.A., Akobundu I.O. & Okonkwo S.N.C., 1991. Effects of Gliricidia sepium (Jacq.) Steud and Leucaena leucocephala (Lam.) de Wit on growth and development of Imperata cylindrica (L.) Raeuschel. Agroforestry Syst., 16, 1-12.
Atta-Krah A.K., 1987. Flowering and seed production of Gliricidia sepium. In: Withington D., Glover N. & Brewbaker J.L., eds. Gliricidia sepium (Jacq.) Walp.: management and improvement. Waimanalo, HI, USA: NFTA (Nitrogen Fixing Tree Association) Special Publication 87-01, 45-50.
Babatoundé S., 2005. Étude et prédiction de la valeur alimentaire de graminées et de légumineuses fourragères en zone tropicale humide du Bénin. Thèse de doctorat: Faculté universitaire des Sciences agronomiques de Gembloux (Belgique).
Barnes P. & Addo-kwafo A., 1996. Evaluation of introduced forage accessions for fodder production at a subhumid site in southern Ghana. Trop. Grassl., 30, 422-425.
Bayer W. & Waters-Bayer A., 1989. Adapting tropical pasture research to the production system: from Australian ranching to African pastoralism. Exp. Agric., 25, 277-289.
Becker M. & Johnson D.E., 1998. Legumes as dry season fallow in upland rice-based systems of West Africa. Biol. Fert. Soils, 27, 358-367.
Bielig L.M., 1997. Chromosome numbers in the forage legume genus, Aeschynomene L. Sabrao J., 29, 33-39.
Bigot S. et al., 2005. Facteurs de la variabilité pluviométrique en Côte d’Ivoire et relations avec certaines modifications environnementales. Sécheresse, 16, 5-13.
Bilgo A. et al., 2007. Chemical and microbial properties of semi-arid tropical soils of short-term fallows in Burkina Faso, West Africa. Biol. Fert. Soils, 43, 313-320.
Bishop H.G., 1992. Aeschynomene americana L. In: ‘t Mannetje L. & Jones R.M., eds. Plant Resources of South-East Asia. No4. Forages. Wageningen, The Netherlands: Pudoc Scientific Publishers.
Bishop H.G., Ludke D.H. & Rutherford M.T., 1985. Glenn jointvetch: a new pasture legume for Queensland coastal areas. Queensland Agric. J., 111, 241-245.
Bishop H.G., Pengelly B.C. & Ludke D.H., 1988. Classification and description of a collection of the legume genus, Aeschynomene. Trop. Grassl., 22, 160-175.
Bogdan A.V., 1977. Tropical pastures and fodder plants (grasses and legumes). London; New York, USA: Longman.
Bosch C.H., Siemonsma J.S., Lemmens R.H. & Oyen L.P., eds, 2002. Plant resources of tropical Africa/Ressources végétales de l’Afrique tropicale. Basic list of species and commodity grouping/Liste de base des espèces et de leurs groupes d’usage. Wageningen, The Netherlands: PROTA Programme.
Bowden B.N., 1963. Studies on Andropogon gayanus Kunth. 1. The use of Andropogon gayanus in agriculture. Emp. J. Exp. Agric., 31, 267-273.
Bray R.A., Tatang I.B. & Schlink A.C., 1993. Yield and quality of Gliricidia sepium accessions at two sites in the tropics. Trop. Grassl., 27, 30-36.
Buckles D., Triomphe B. & Sain G., 1998. Cover crops in hillside agriculture. Farmer innovation with Mucuna. Ottawa: IDRC and CIMMYT.
Buldgen A. & Dieng A., eds., 1997. Andropogon gayanus var. ‘bisquamulatus’. Une culture fourragère pour les régions tropicales. Gembloux, Belgique: Les Presses agronomiques de Gembloux.
Buldgen A. et al., 2001. Production and nutritive value of grasses cultivated in coastal area of Benin. Trop. Grassl., 35, 43-47.
Cameron D.G., 1985. Tropical and subtropical legumes. 5. Siratro pasture (Macroptilium atropurpureum): the most widely planted subtropical legume. Queensland Agric. J., 111, 45-49.
CARDER (Centre d’Action Régionale de Développement Rural-Borgou), 2005. Rapport annuel d’activité. Borgou, Bénin: CARDER.
Carr P.M., Poland W.W. & Tisor L.J., 2005. Natural reseeding by forage legumes following wheat in western North Dakota. Agron. J., 97, 1270-1277.
Carsky R.J. et al., 1998. Mucuna: herbaceous cover legume with potential for multiple uses. Resource and crop management research monograph. 25. Ibadan, Nigeria: International Institute of Tropical Agriculture (IITA).
C.P.C.S. (Commission de Pédologie et Cartographie des Sols), 1967. Classification française des sols. Travaux C.P.C.S. Grignon, France: ENSA.
Clem R.L., 2004. Animal production from legume-based ley pastures in southeastern Queensland. In: Whitbread A.M. & Pengelly B.C., eds. Tropical legumes for sustainable farming systems in Southern Africa and Australia. Proceedings no 115. Canberra: ACIAR, 136-144.
Cobbina J. & Atta-Krah A.N., 1992. Gliricidia sepium accessions on a tropical alfisol in Nigeria. Trop. Grassl., 26, 248-254.
Cook B.G. et al., 2005. The production of tropical forages: an alternative selection tool. Http://www.tropicalforages.info, (15/02/08).
Cullen B.R. & Hill J.O., 2006. A survey of the use of lucerne, butterfly pea and lablab in ley pastures in the mixed-farming systems of northern Australia. Trop. Grassl., 40, 24-32.
Dalal R.C. & Mayer R.J., 1986. Long-term trends in fertility of soils under continuous cultivation and cereal cropping in southern Queensland. 1. Overall changes in soil properties and trends in winter cereal yields. Aust. J. Soil Res., 24, 265-279.
Dalal R.C., Strong W.M., Weston E.J. & Gaffney J., 1991. Sustaining multiple production systems. 2. Soil fertility decline and restoration of cropping lands in sub-tropical Queensland. Trop. Grassl., 25, 173-180.
Dennison E.B., 1959. The maintenance of soil fertility in the southern Guinea Savanna zone of southern Nigeria. Trop. Agric., 36, 330-336.
Dommergues Y., Duhoux E. & Diem H.G., 1999. Les arbres fixateurs d’azote. Montpellier, France: Éditions Espaces 34.
Dzowela B.H., Wandera P.F., Were J. & Mohammed- Saleem M.A., 1998. Leucaena in smallholder farming in Africa: challenge for development. In: Proceedings of a workshop held in Hanoi, Vietnam, 9-14 February, 311-318.
Edye L.A. & Maass B.L., 1997. Recent advances in studies of anthracnose of Stylosanthes. I. The biogeography of Stylosanthes hamata, Stylosanthes scabra and Stylosanthes seabrana. Trop. Grassl., 31, 417-423.
Eilittä M. et al., 2003. Increasing Mucuna’s potential as a food and feed crop. Proceedings of an international workshop held September 23-26, 2002, in Mombasa, Kenya. Trop. Subtrop. Agroecosystem, 1(special issue), 1-343, http://www.uady.mx/sitios/veterina/servicios/journal/2002-2-3/indice-1(2-3).html, (20/01/08).
Ella A., Jacobsen C., Stür W.W. & Blair G., 1989. Effect of plant density and cutting frequency on the productivity of four tree legumes. Trop. Grassl., 23, 28-34.
English B.H., 1999. Macroptilium atropurpureum in Australia. In: Loch D.S. & Ferguson J.E., eds. Forage seed production. 2: Tropical and subtropical species. Wallingford, UK: CABI Publishing, 407-412.
Falade J.A., 1975. The effect of phosphorus on the growth and mineral composition of five tropical grasses. East Afr. Agric. For. J., 40, 342-350.
Fantz P.R., 1996. Taxonomic notes on the Centrosema pubescens Bentham complex in Central America (Leguminosae: Phaseoleae: Clitoriinae). Sida, 17, 321-332.
FAO (Food and Agriculture Organisation), 1965. Guidelines for soil description. Paris: FAO Flat.
Floret C. & Pontannier R., 1999. La jachère en Afrique tropicale. 1: Les actes du congrès. Rôles, aménagement, alternatives. In: Actes du séminaire international, Dakar, Sénégal, avril 1999.
Gbangboché A.B., 2005. Performances zootechniques et paramètres génétiques des ovins Djallonké au Bénin. Thèse de doctorat: Université de Liège (Belgique).
Geerts P., Buldgen A., Diallo T. & Dieng A., 1998. Drought resistance by six Senegalese local strains of Andropogon gayanus var. ‘biscumulatus’ through osmoregulation. Trop. Grassl., 32, 235-242.
Gichuru M.P. & Kang B.T., 1989. Calliandra calothyrsus (Meissn.) in an alley cropping system with sequentially cropped maize and cowpea in southwestern Nigeria. Agroforestry Syst., 9, 191-203.
Gillett J.B., Polhill B.A. & Verdcourt B., 1971. Leguminosae, Papilionoideae. In: Milne-Redhead E. & Polhill R.M., eds. Flora of tropical East Africa. London: Crown agents for overseas governments and administrations.
Gobius N.R. et al., 2001. Seed yield and its components of Brachiaria decumbens cv. ‘Basilis’, Digitaria milanjiana cv. ‘Jarra’ and Andropogon gayanus cv. ‘Kent’ in north-east Thailand under different rates of nitrogen application. Trop. Grassl., 35, 26-33.
Grof B., 1968. Viability of seed of Brachiaria decumbens. Queensland J. Agric. Anim. Sci., 25, 149-152.
Haggar R.J., 1966. The production of seed from Andropogon gayanus. In: Proceedings of the International Seed Testing Association (I.S.T.A.), 31, 251-259.
Hakiza J.J., Lazier J.R. & Sayers A.R., 1987. Characterization and evaluation of forages legumes in Ethiopia: preliminary examination of variation between accessions of Stylosanthes fruticosa (Retz.) Alston. In: Dzowela B.H., ed. African forage plant genetic resources, evaluation of forage germplasm and extensive livestock production systems. Proceedings of the 3rd PANESA (Pasture Network for Eastern and Southern Africa) workshop held at Arusha, Tanzania, 27-30 April 1987, 174-191.
Hall T.J. & Glatzle A., 2004. Cattle production from Stylosanthes pastures. In: Chakraborty S., ed. High yielding anthracnose-resistant Stylosanthes for agricultural systems. Monograph no111. Canberra: ACIAR.
Hare M.D. & Phaikew C., 1997. Forage seed production in northeast Thailand. In: Loch D.S. & Ferguson J.E., eds. Forage seed production. 2: Tropical and subtropical species. Wallingford, UK: CABI Publishing, 435-443.
Hare M.D., Tatsapong P. & Saipraset K., 2007. Seed production of two brachiaria hybrid cultivars in northeast Thailand. 1. Method and time of planting. Trop. Grassl., 41, 26-34.
Häussler K., Rao I.M., Schultze-Kraft R. & Marschner H., 2006. Shoot and root growth of two tropical grasses, Brachiaria ruziziensis and Brachiaria dictyoneura, as influenced by aluminium toxicity and phosphorus deficiency in a sandy loam oxisol of the eastern plains of Colombia. Trop. Grassl., 40, 213-221.
Hegarty M.P., Court R.D., Christie G.S. & Lee C.D., 1976. Mimosine in Leucaena leucocephala is metabolised to a goitrogen in ruminants. Aust. Vet. J., 52, 490-495.
Hodges E.M. et al., 1982. Production and utilization of the tropical legume Aeschynomene (Aeschynomene americana L.). Circular S-290. Gainesville, FL., USA: Institute of Food and Agricultural Sciences, University of Florida.
Hopkinson J.M. et al., 1996. Reproductive physiology, seed production and seed quality of Brachiaria. In: Miles J.W., Maass B.L., Valle C.B. & do Kumble V., eds. Brachiaria: biology, agronomy, and improvement. Cali, Colombia: CIAT Publication no259, 124-140.
Humphreys L.R., 1994. Tropical forages: their role in sustainable agriculture. Essex, UK: Longman.
Jones R.J., 1979a. The value of Leucaena leucocephala as a feed for ruminants in the tropics. World Anim. Rev., 31, 13-23.
Jones C.A., 1979b. The potential of Andropogon gayanus Kunth in the oxisol and ultisol savannas of Tropical America. Herb. Abstr., 19, 1-8.
Jones R.J., 1986. Leucaena: international experience. Trop. Grassl., 20, 83-85.
Jones R.J. & Jones R.M., 1982. Observations on the persistence and potential for beef production of pastures based on Trifolium semipilosum and Leucaena leucocephala in sub-tropical coastal Queensland. Trop. Grassl., 16, 24-29.
Jones R.M. & Bunch G.A., 2003a. Experiences with farm pastures at the former CSIRO Samford Research Station, southeast Queensland, and how these relate to results from 40 years of research. Trop. Grassl., 37, 151-164.
Jones R.M. & Jones R.J., 2003b. Effect of stocking rates on animal gain, pasture yield and composition, and soil properties from setaria-nitrogen and setaria-legume pastures in coastal southeast Queensland. Trop. Grassl., 37, 65-83.
Kaminski P.E., Schifino-Wittmann M.T. & Paim N.R., 2005. Growth and survival of a range of Leucaena species in southern Brazil. Trop. Grassl., 39, 1-8.
Klein H.D. & César J., 1999. Plantes fourragères et fertilité du sol. In: Robertge G. & Toutain B. Cultures fourragères tropicales. Montpellier, France: CIRAD, 321-357.
Koutika L.S., Sanginga N., Vanlauwe B. & Weise S., 2002. Chemical properties of soil organic matter assessment under fallow systems in the forest margins benchmark. Soil Biol. Biochem., 34, 757-765.
Kretschmer A.E. & Bullock R.C., 1980. Aeschynomene spp.: distribution and potential use. In: Proceedings of the Soil and Crop Science Society of Florida, 39, 145-152.
Lalljee B., Facknath S. & Osman A., 1998. Improvement of soil properties under long-term Leucaena leucocephala. In: Shelton H.M., Gutteridge R.C., Mulen B.F & Bray R.A., eds. Leucaena: adaptation, quality and farming systems. Proceedings no86. Canberra: ACIAR, 178-180.
Mandret G. & Noirot M., 1999. Production de semences fourragères tropicales. In: Roberge G. & Toutain B., eds. Cultures fourragères tropicales. Montpellier, France: CIRAD, 189-213.
Mapiye C., Mwale M. & Mupangwa J.F., 2007. Utilization of ley legumes as livestock feed in Zimbabwé. Trop. Grassl., 41, 84-91.
Mapongmetsem P.M., Duguma B., Nkongmeneck B.A. & Selegny E., 1999. Germination des semences, développement et croissance de quelques essences locales en zone forestière. Tropicultura, 17, 175-179.
McIntire J., Bourzat D. & Pingali P., 1992. Crop livestock interaction in sub-Saharan Africa. Washington, DC, USA: World Bank.
McKeon G.M. & Mott J.J., 1984. Seed biology of Stylosanthes. In: Stace H.M. & Edye L.A., eds. The biology and agronomy of Stylosanthes. North Ryde, Australia: Academic Press, 311-332.
McNeill D.M., Osborne N., Komolong M. & Nankervis D., 1998. Condensed tannins in the genus Leucaena and their nutritional significance for ruminants. In: Shelton H. M., Gutteridge R.C., Mullin B.F. & Bray R.A., eds. Leucaena: adaptation, quality and farming systems. Proceedings no86. Canberra: ACIAR, 205-214.
Merkel U. et al., 2000. Characterization of a collection of Aeschynomene histrix in subhumid Nigeria. J. Agric. Sci., 134, 293-304.
Messager J.L., 1984. Notes sur les modalités techniques d’installation des pâturages de Panicum. Rev. Élev. Méd. Vét. Pays Trop., 37, 336-340.
Middleton C.H. & McCosker T.H., 1975. Makueni, a new guinea grass for North Queensland. Queensland Agric. J., 101, 351-355.
Miles J.W., Maass B.L., Valle C.B. & do Kumble V., eds., 1996. Brachiaria: biology, agronomy, and improvement. Cali, Colombia: CIAT Publication no259.
Mitja D., Leal N. & Topall O., 1998. Rehabilitation of degraded Amazon pastures. Cases of Andropogon gayanus Kunth (Maraba, Para, Brazil). Rev. Écol. Terre Vie, 53, 39-57.
Miyagi E., 1980. The effect of planting density on yield of napier grass (Pennisetum purpureum Schumach). Sci. Bul. Coll. Agric., 27, 293-301.
Mohamed-Saleem A.M., 1986. Integration of forage legumes into the cropping systems of Nigeria’s sub-humid zone. In: Kaufmann V.R., Chater S. & Blench R., eds. Livestock systems research in Nigeria’s sub-humid zone. Proceedings of the second ILCA/NAPRI Symposium held in Kaduna-Nigeria, 29 October-2 November 1984, Addis- Abeba, Ethiopia, 305-325.
Mugwe J.N., O’Neil M.K. & Mugendi N.D., 1998. Leuceuna leucocephala and Calliandra calothyrsus for soil productivity improvement in sub-humid Highlands of Kenya. In: Proceedings of a workshop held on February 9-14 in Hanoi, Vietnam, 181-184.
Muhr L., 1998. Potential of forage legumes on fallow land for enhanced forage production and soil fertility in crop-livestock farming systems of subhumid West Africa. Thesis: University Hohenheim (Germany).
Muhr L., Peters M., Tarawali S.A. & Schultze-Kraft R., 1999. Forage legumes for improved fallows in agropastoral systems of subhumid West Africa. III. Nutrient import and export by forage legumes and their rotational effects on subsequent maize. Trop. Grassl., 33, 245-256.
Muhr L., Tarawali S.A., Peters M. & Schultze-Kraft R., 2002. Soil mineral N dynamics and maize grain yields following Centrosema macrocarpum and Stylosanthes guianensis: effects of different rotations and varying levels of fertilizer. Field Crops Res., 78, 197-209.
Muia J.M.K., Tamminga S., Mbugua P.N. & Kariuki J.N., 1999. Optimal stage of maturity for feeding napier grass (Pennisetum purpureum) to dairy cows in Kenya. Trop. Grassl., 33, 182-190.
Mullen B.F., Shelton H.M. & Dalzell S.A., 2005. Leucaena in northern Australia: a forage tree legume success story. Trop. Grassl., 39, 226.
Muyekho F.N., Mose L. & Cheruiyot D.T.,2003.Development and transfer of forage production technologies for smallholder dairying: case studies of participatory evaluation of species and methods of establishment in western Kenya. Trop. Grassl., 37, 251-256.
Nikiema A., 2005. Agroforestry parkland species diversity: uses and management in semi-arid west Africa (Burkina Faso). Thesis: Wageningen University (The Netherlands).
Noirot M. et al., 1986. La production grainière des nouvelles variétés de Panicum maximum sélectionnées en Côte d’Ivoire. Fourrages, 106, 11-18.
Nye P.H. & Greenland D.J., 1960. The soil under shifting cultivation. Technical communication no51. Harpenden, UK: Commonwealth Bureau of Soils; Farnham Royal, UK: Commonwealth Agricultural Bureaux.
Olanite J.A., Tarawali S.A. & Akan’ova M.E., 2004. Biomass yield, quality and acceptability of selected grass-legume mixtures in the moist savanna of West Africa. Trop. Grassl., 38, 117-128.
Omokanye A.T., 2001. Forage yield and chemical composition of centro (Centrosema pubescens) in the year of establishment at Shika, Nigeria. Trop. Grassl., 36, 53-57.
Ong C.K. & Daniel J.N., 1990. Le pois d’Angole. Agroforesterie Aujourd’hui, 2, 4-7.
Peake D.C.I., Myers R.J.K. & Henzell E.F., 1990. Sown pasture production in relation to nitrogen fertiliser and rainfall in Southern Queensland. Trop. Grassl., 24, 291-298.
Pengelly B.C., Clem R.L. & Whitbread A.M., 2004. The role of Stylosanthes spp. in mixed crop-livestock systems in Africa and Australia. In: Chakraborty S., ed. High yielding anthracnose-resistant Stylosanthes for agricultural systems. Monograph no111. Canberra: ACIAR.
Peters M. et al., 2001. The role of forages in reducing poverty and degradation of natural resources in tropical production systems. Network paper n°117. London: Agricultural Research and Extension Network, http://www.odi.org.uk/agren/publist.html, (15/02/08).
Peters M. & Lascano C.E., 2003. Forage technology adoption: linking on-station research with participatory methods. Trop. Grassl., 37, 197-203.
Phaikaew C. & Pholsen P., 1993. Ruzi grass (Brachiaria ruziziensis) seed production and research in Thailand. In: Chen C.P. & Satjipanon C., eds. Strategies for suitable forage-based livestock production in Southeast Asia. Proceedings of the 3rd Meeting of the regional working group on grazing and feed resources of Southeast Asia, Department of Livestock Development, Bangkok, Thailand, 165-173.
Piggin C.M. & Nulik J., 2005. Leucaena: sustainable crop and livestock production systems in Nusa Tenggara Timur province, Indonesia. Trop. Grassl., 39, 218.
Rao I.M., Boddey R.M. & Thomas R.J., 1996. Nutrient cycling and environmental impact of brachiaria pastures. In: Miles J.W., Maass B.L., Valle C.B. & do Kumble V., eds. Brachiaria: biology, agronomy and improvement. Cali, Colombia: CIAT, 72-86.
Rethman N., 2000. Grass, grass-legume or legume leys: a south African experience. Trop. Grassl., 34, 177-180.
Roberge G. & Toutain B., 1999. Choix des plantes fourragères. In: Roberge G. & Toutain B., eds. Cultures fourragères tropicales. Montpellier, France: CIRAD, 147-184.
Rudd V.E., 1955. The American species of Aeschynomene. Contributions from the United States. Natl Herb., 32, 23-30.
Russell J.S., 1991. Likely climatic changes and their impact on the northern pastoral industry. Trop. Grassl., 25, 211-218.
Saïdou A., 2006. Converting strategies by farmers and scientists to improve soil fertility and enhance crop- production in Benin. Thesis: Wageningen University (The Netherlands).
Sanginga N. et al., 1996. Nodulation and estimation of symbiotic nitrogen fixation by herbaceous and shrub legumes in Guinea savanna in Nigeria. Biol. Fert. Soils, 23, 441-448.
Santos P.M., Corsi M., Pedreira C.G.S. & Lima C.G., 2006. Tiller cohort development and digestibility in Tanzania guinea grass (Panicum maximum cv. ‘Tanzania’) under three levels of grazing intensity. Trop. Grassl., 40, 84-93.
Schultze-Kraft R. & Clements R.J., eds., 1990. Centrosema: biology, agronomy and utilization. Cali, Colombia: CIAT.
Schultze-Kraft R. & Peters M., 1997. Tropical legumes in agricultural production and resource management: an overview. Giessener Beitr. Entwicklungsforschung, 24, 1-17.
Shehu Y., Alhassan W.S. & Phillips C.J.C., 1997. The effect of intercropping maize with Stylosanthes hamata at different row spacings on grain and fodder yields and chemical composition. Trop. Grassl., 31, 227-231.
Sidi-Imorou H., 2002. Étude du comportement alimentaire des moutons Djallonké et de leurs performances à l’engraissement sur les cultures fourragères en mélange. Thèse de doctorat: Université d’Abomey-Calavi (Bénin).
Sinsin B., Oloulotan S. & Oumorou M., 1989. Les pâturages de saison sèche de la zone soudanienne du nord-est du Bénin. Rev. Élev. Méd. Vét. Pays Trop., 42, 283-288.
Skerman P.J. et al., 1988. Tropical forage legumes. FAO plant production and protection series no2. Roma: FAO.
Somé N.A., Hien V. & Ouédraogo S.J., 2006. Amélioration du statut organique d’un sol ferrugineux tropical lessivé sous jachère artificielle à Andropogon spp. au Burkina Faso: effet des traitements sur la chimie du sol. Tropicultura, 24, 200-207.
Somé N.A., Traoré K., Traoré O. & Tassembedo M., 2007. Potentiel des jachères artificielles à Andropogon spp. dans l’amélioration des propriétés chimiques et biologiques des sols en zone soudanienne (Burkina Faso). Biotechnol. Agron. Soc. Environ., 11, 245-252.
Stace H.M. & Edye L.A., eds., 1984. The biology and agronomy of Stylosanthes. North Ryde, Australia: Academic Press.
Sultan B. & Janicot S., 2004. La variation climatique en Afrique de l’Ouest aux échelles saisonnière et intra- saisonnière. 1. Mise en place de la mousson et variabilité intra-saisonnière de la convection. Sécheresse, 15, 321-330.
Mannetje L., 1992. Pennisetum purpureum Schumach. In: ‘t Mannetje L. & Jones R.M., eds. Plant resources of Southeast Asia. 4: Forages. Wageningen, The Netherlands: Pudoc Scientific Publishers, 191-192.
Tarawali G. & Mohamed-Saleem M.A., 1995. The role of forage legume fallows in supplying improved feed and recycling nitrogen in subhumid Nigeria. In: Tarawali G. et al., eds. Livestock and sustainable nutrient cycling in mixed farming systems of sub-Saharan Africa. 2: Technical papers. Proceedings, Addis-Abeba, Ethiopia, 22-26 November 1993, 263-276.
Toléba S.S. et al., 2001. Étude comparative de deux espèces fourragères (Panicum maximum local et Brachiaria ruziziensis) complémentées par des graines de coton sur les performances pondérales des ovins Djallonké. Ann. Sci. Agron. Bénin, 2, 193-208.
Toledo J.M., Vera R., Lascano C. & Lenne J.M., eds., 1990. Andropogon gayanus Kunth: a grass for tropical acid soils. Publication no90. Cali, Colombia: CIAT.
Traoré K., Ganry F., Olivier R. & Gigou J., 2004. Litter production and soil fertility in a Vitellaria paradoxa parkland in a catena in Southern Mali. Arid Land Res. Manage., 18, 359-368.
Tudsri S., Jorgensen S.T., Riddachi P. & Pookpakdi A., 2002. Effect of cutting height and dry season closing date on yield and quality of five napier grass cultivars in Thailand. Trop. Grassl., 36, 248-252.
Valarini M.J. & Possenti R.A., 2006. Research note: nutritive value of a range of tropical forage legumes. Trop. Grassl., 40, 183-187.
Vallis I. & Gardener C.J., 1984. Nitrogen inputs into agricultural systems by Stylosanthes. In: Stace H. M. & Edye L.A., eds. The biology and agronomy of Stylosanthes. North Ryde, Australia: Academic Press, 359-379.
Vandenbeldt J.R., 1988. Cajanus cajan: it’s more than just a pulse. NFT Highlights, NFTA 88(6), 1-2.
Walmsley D., Sargeant V.A.L. & Dookeran M., 1978. Effect of fertilizers on growth and composition of elephant grass (Pennisetum purpureum) in Tobago, West Indies. Trop. Agric., 25, 329-334.
Wandera F.P., Karachi M., Mangope S. & Lefofe B.M., 2005. Leucaena production in Botswana. Trop. Grassl., 39, 229.
Weber G. et al., 1995. Striga hermonthica (Del.) Benth. in the cropping systems of the Northern Guinea savanna. Resource and Crop Management Division. Vol. 19. Ibadan, Nigeria: International Institute of Tropical Agriculture (IITA).
Whitbread A.M., Jiri O. & Maasdorp B., 2004. The effect of managing improved fallows of Mucuna pruriens on maize production and soil carbon and nitrogen dynamics in sub-humid Zimbabwe. Nutr. Cycling Agroecosystem, 69, 59-71.
Youssouf I. & Lawani M., 2002. Les sols béninois: classification dans la référence mondiale. In: Quatorzième réunion du sous-comité ouest et centre africain de corrélation des sols pour la mise en valeur des terres. 9-13 octobre 2000, Bénin, Abomey-Calavi. Rome: FAO.