lochkovian; trilete spores; cryptospores; palaeophytogeography; palacoecology; Saudi Arabia
Abstract :
[en] Palynological samples from two boreholes drilled in the eastern part of Saudi Arabia contain a rich assemblage of cryptospores and trilete spores. They are dated as most probably Lochkovian in age. Intriguingly, the cryptospores are very similar to those from a coeval continental palynomorph assemblage from the Old Red Sandstone Continent, whereas the trilete spores show a strong affinity with miospore assemblages from Western Gondwana. It is suggested that cryptospore-producing plants inhabited confined, damp biotopes and had a wide range of climatic tolerance. On the otherhand, trilete spore-producing plants inhabited a wider variety of biotopes but were more sensitive to climatic variations. The high degree of similarity between Old Red Sandstone and Saudi Arabian cryptospore assemblages favours palaeogeographic reconstructions where there is close proximity between Western Gondwana and Euramerica and/or a land connection between these palaeoplates. Climatic differences between the Old Red Sandstone Continent and the northern part of Western Gondwana can be invoked to explain the variations in the trilete spore assemblages. (c) 2007 Elsevier B.V. All rights reserved.
Abu-Ali M., and Littke R. Paleozoic petroleum systems of Saudi Arabia: a basin modeling approach. GeoArabia 10 3 (2005) 131-168
Al-Ameri, T.K., 1980. Palynology, biostratigraphy and palaeoecology of subsurface Mid-Palaeozoic strata from the Ghadames Basin, Libya. PhD Thesis. University of London, UK.
Al-Hajri S., and Owens B. Sub-surface palynostratigraphy of the Paleozoic of Saudi Arabia. In: Al-Hajri S., and Owens B. (Eds). Stratigraphic Palynology of the Paleozoic of Saudi Arabia. Special GeoArabia Publication vol. 1 (2000), Gulf Petrolink, Manama, Bahrain 10-17
Al-Hajri S., and Paris F. Age and paleoenvironment of the Sharawra Member (Silurian of northwestern Saudi Arabia). Geobios 31 1 (1998) 3-12
Al-Hajri S., Filatoff J., Wender L.E., and Norton A.K. Stratigraphy and operational palynology of the Devonian System in Saudi Arabia. GeoArabia 4 1 (1999) 53-68
Al-Husseini M., and Mathews R.K. Arabian orbital stratigraphy: periodic second-order sequence boundaries?. GeoArabia 10 2 (2005) 165-184
Al-Laboun A.A., and Walthall B.H. The Devonian of the Arabian Peninsula. In: McMillan N.J., Embry A.F., and Glass D.J. (Eds). Devonian of the World: Proceedings of the 2nd International Symposium on the Devonian System, Calgary Canada, 1987: Regional Synthesis vol. 1 (1988), Canadian Society of Petroleum Geologists Publication 569-577
Aoudeh S.M., and Al-Hajri S.A. Regional distribution and chronostratigraphy of the Qusaiba Member of the Qalibah Formation in the Nafud Basin, northwestern Saudi Arabia. In: Al-Husseini M.I. (Ed). The Middle East Petroleum Geosciences, Geo '94 vol. 1 (1994), Gulf PetroLink, Manama, Bahrain 143-154
Arkhangelskaya A.D. Plant spores from some Lower Devonian sections of the western regions of the Russian Plate. In: Byvsheva T.V. (Ed). Palynological Research in the Proterozoic and Phanerozoic of Oil and Gas Bearing Regions of USSR. Trudy Vsesoluznogo Naucho-Issledovate skogo Geologoazvedochnogo Neftianogo Instituta, Kama Branch (1980) 26-46 (in Russian)
Beck J.H., and Strother. Silurian spores and cryptospores from the Arisaig Group, Nova Scotia, Canada. Palynology 25 (2001) 127-177
Beju D. Quelques spores, acritarches et chitinozoaires d'âge Dévonien inférieur de la plate-forme moesienne (Roumanie). Review of Palaeobotany and Palynology 5 (1967) 39-49
Boucot A.J., McClure H.A., Alverez F., Ross J.R.P., Taylor D.W., Struve W., Savage N.N., and Turner S. New Devonian fossils from Saudi Arabia and their biogeographical affinities. Senckenbergiana Lethaea 69 5/6 (1989) 535-597
Breuer P., Al-Ghazi A., Filatoff J., Higgs T., Steemans P., and Wellman C. Stratigraphic palynology of Devonian boreholes from northern Saudi Arabia. Carnets de Géologie/Notebooks on Geology, Brest, Memoir 2005/02 (2005) Abstract 01 (CG2005_M02/01)
Breuer P., Stricanne L., and Steemans P. Morphological variation within a miospore population belonging to the genus Emphanisporites McGregor 1961, Lower Devonian, "la Gileppe" outcrop, Belgium. Geological Magazine 142 3 (2005) 241-253
Buatois L.A., and Mángano M.G. Animal-substrate interactions in freshwater environments: applications of ichnology in facies and sequence stratigraphic analysis of fluviolacustrine successions. In: McIlroy D. (Ed). The Application of Ichnology and Stratigraphic Analysis. Geological Society, London, Special Publications vol. 228 (2004) 311-333
Burgess N.D. Silurian cryptospores and miospores from the type Llandovery area, south-west Wales. Palaeontology 34 3 (1991) 575-599
Burgess N.D., and Richardson J.B. Late Wenlock to early Pridoli cryptospores and miospores from south and southwest Wales, Great Britain. Palaeontographica 236(Abt. B) (1995) 1-44
Cocks L.R.M., and Torsvik T.H. Earth geography from 500 to 400 million years ago: a faunal and palaeomagnetic review. Journal of the Geological Society (London) 159 (2002) 631-644
Cramer F.H. Palynomorphs from the Siluro-Devonian boundary in N. W. Spain. Notas y Comunicaciones del Instituto Geologico y Minero de Espana 85I (1966) 71-82
Cramer F.H. Microplankton from three Palaeozoic formations in the Province of León (NW Spain). Leidse Geol. Meded. 30 (1964) 253-361
Cramer F.H., and Díez M.D.C.R. North American Silurian palynofacies and their spatial arrangement. Acritarchs. Palaeontographica, abt. B 138 5/6 (1972) 107-180
Cramer F.H., and Rodríguez R.M. Robledo and Arroyacas Formation (Arroyo de las Arroyacas, Province of Palencia, Spain) palynologically dated as Late Silurian. Breviora Geologica Asturica 21 1 (1977) 2-4
Cramer F.H., Díez M.D.C.R., Rodríguez R.M., and Fombella M.A. Acritarcos de la Formación San Pedro (Silúrico superior) de Torrestio, Provincia de León, España. Revista Española Micropaleontologia VIII 3 (1976) 439-452
D'Erceville M.A. Les spores des formations siluro-dévoniennes, de la coupe de Saint-Pierre-sur-Erve, synclinorium médian armoricain. Palaeontographica 171 (1979) 79-121 (B)
Edwards D., and Richardson J.B. Silurian to Lower Devonian plant assemblages from the Anglo-Welsh Basin: a palaeobotanical and palynological synthesis. Geological Journal 39 (2004) 375-402
Edwards D., and Wellman C.H. Embryophytes on land: the Ordovician to Lochkovian (Lower Devonian) Record. 3-28. In: Gensel P.G., and Edwards D. (Eds). Plants Invade the Land (2001), Columbia University Press, New York 204 pp.
Evans D.S., Lathon R.B., Senalp M., and Connally T.C. Stratigraphy of the Wajid Sandstone of southwestern Saudi Arabia. Society of Petroleum Engineers, Publication, SPE 21449 (1991) 947-960
Gao-Liando. Devonian spore assemblages of China. Review of Palaeobotany and Palynology 34 (1981) 11-23
Gessa S., Lardeux H., Gourvennec R., Becq-Giraudon J.-F., Janjou D., Halawani M.A., Robelin C., Al Muallem M., and Brosse J.-M. A new species of tentaculitids from the Qalibah Formation, NW Saudi Arabia. Revue de Micropaléontologie 8 3 (1994) 201-208
Gradstein F.M., Ogg J.G., and Smith A.G. A Geological Time Scale (2004), Cambridge University Press 589 pp.
Gray J. The microfossil record of early land plants; advances in understanding of early terrestrialization, 1970-1984. In: Chaloner W.G., and Lawson J.D. (Eds). Evolution and Environment in the Late Silurian and Early Devonian. Philosophical Transactions of the Royal Society, London (1985) 167-195
Hance L., Dejonghe L., Fairon-Demaret M., and Steemans P. La Formation de Pépinster dans le Synclinorium de Verviers, entre Pépinster et Eupen (Belgique) - Contexte structural et stratigraphique. Annales de la Société Géologique de Belgique 117 1 (1996) 75-93
Haq B.U., and Al-Qahtani A.M. Phanerozoic cycles of sea-level change on the Arabian Platform. GeoArabia 10 2 (2005) 127-160
Jaglin J.C., and Paris F. Biostratigraphy, biodiversity and palaeogeography of Late Silurian chitinozoans from A1-61 borehole (north-western Libya). In: Steemans P., Servais T., and Streel M. (Eds). Palaeozoic Palynology: A Special Issue in Honour of Dr. Stanislas Loboziak. Review of Palaeobotany and Palynology, Amsterdam (2002) 335-358
Janjou, D., Halawani, M., Al-Muallem, M.S., Robelin, C., Brosse. J.-M., Courbouleix, S., Dagain, J., Genna, A., Razin, P., Roobol, M.J., Shorbaji, Hassan and Wyns, R., 1997a. Explanatory notes to the geological map of the Al Qalibah Quadrangle, Kingdom of Saudi Arabia. Geoscience Map G-135, scale 1:250,000, sheet 28C. Deputy Ministry for Mineral Resources, Ministry of Petroleum and Mineral Resources, Kingdom of Saudi Arabia, 44 pp.
Janjou, D., Halawani, M.A., Brosse, J.-M., Al-Muallem, M.S., Becq-Giraudon, J.F., Dagin, J., Genna, A., Razin, M.J., RooBol, M.J., Shorbaji, H., Wyns, R., 1997b. Explanatory notes to the geological map of the Tabuk Quadrangle, Kingdom of Saudi Arabia. Geoscience Map G-137, scale 1:250,000, sheet 28B. Deputy Ministry for Mineral Resources, Ministry of Petroleum and Mineral Resources, Kingdom of Saudi Arabia, 49 pp.
Jardiné S., and Yapaudjian L. Lithostratigraphie et palynologie du Dévonien-Gothlandien gréseux du Bassin de Polignac (Sahara). Revue de l'Institut Français du Pétrole 23 4 (1968) 439-469
Johnson N.G. Early Silurian palynomorphs from the Tuscarora Formation in Central Pennsylvania and their paleobotanical and geological significance. Review of Palaeobotany and Palynology 45 3/4 (1985) 307-360
Juan C., Emerson B.C., Oromi P., and Hewitt G.M. Colonization and diversification: towards a phylogoegraphic synthesis for the Canary Islands. Tree 13 3 (2000) 104-109
Konert G., Afifi A.M., Al-Hajri S.A., and Droste H.J. Paleozoic stratigraphy and hydrocarbon habitat of the Arabian Plate. GeoArabia 6 3 (2001) 407-441
Lavender K., and Wellman C.H. Lower Devonian spore assemblages from the Arbuthnott Group at Canterland Den in the Midland Valley of Scotland. In: Steemans P., Servais T., and Streel M. (Eds). Palaeozoic Palynology: A Special Issue in Honour of Dr. Stanislas Loboziak. Review of Palaeobotany and Palynology, Amsterdam (2002) 157-180
Le Hérissé A. Les spores du Dévonien inférieur du Synclinorium de Laval (Massif Armoricain). Palaeontographica Abteilung B Paläophytologie 188 (1983) 1-81
Le Hérissé A. Paleoecology, biostratigraphy and biogeoraphy of Late Silurian to Early Devonian acritarchs and prasinophycean phycomata in well A1-61, Western Libya, North Africa. In: Steemans P., Servais T., and Streel M. (Eds). Palaeozoic Palynology: A Special Issue in Honour of Dr. Stanislas Loboziak. Review of Palaeobotany and Palynology, Amsterdam (2002) 359-395
Lele K.M., and Streel M. Middle Devonian (Givetian) plant micofossils from Goé (Belgium). Annales de la Société Géologique de Belgique 92 1 (1969) 89-121
Loboziak S., and Streel M. Middle-Upper Devonian miospores from the Ghadamis Basin (Tunisia-Libya): systematics and stratigraphy. Review of Palaeobotany and Palynology 58 (1989) 173-196
Loboziak S., and Streel M. Late Lower and Middle Devonian miospores from Saudi Arabia. Review of Palaeobotany and Palynology 89 1-2 (1995) 105-113
Lozej, G.P., 1983. Geological and geochemical reconnaissance exploration of the cover rocks in northwestern Hijaz - initial results and recommendations. Deputy Ministry for Mineral Resources, Ministry of Petroleum and Mineral Resources, Kingdom of Saudi Arabia, Open-File Report RF-OF-03-2, 135 pp.
McDaniel S.F., and Shaw A.J. Phylogeographic structure and cryptic speciation in the trans-Antarctic moss Pyrrhobryum mnioides. Evolution 57 (2003) 205-215
McGillivray J.G., and Husseini M.I. The Paleozoic petroleum geology of central Arabia. Bulletin of the American Association of Petroleum Geologists 76 10 (1992) 1473-1490
McGregor D.C. Lower and Middle Devonian spores of Eastern Gaspé, Canada. Palaeontographica Abteilung B Paläophytologie 142 (1973) 1-77
McGregor D.C. Early Devonian spores from central Ellesmere Island, Canadian Artic. Canadian Journal of Earth Sciences 11 (1974) 70-78
McGregor D.C. Lower and Middle Devonian spores of Eastern Gaspe, Canada: II. Biostratigraphy. Palaeontographica Abteilung B Paläophytologie 163 (1977) 111-142
McGregor D.C. Late Silurian and Devonian spores from Bolivia. Academia Nacionaldde Ciencias, Córdoba, Miscelanea 69 (1984) 57
McGregor D.C., and Camfield M. Upper Silurian? to Middle Devonian spores of the Moose River Basin, Ontario. Geological Survey of Canada Bulletin 263 (1976) 1-63
McGregor D.C., and Narbonne G.M. Upper Silurian trilete spores and other microfossils from the Read Bay Formation, Cornwallis Island, Canadian Arctic. Canadian Journal of Earth Sciences 15 (1978) 1292-1303
Mogensen G.S. The biological significance of morphological chracters in bryophytes: the spore. The Bryologist 84 (1981) 187-207
Moore P.D., Webb J.A., and Collinson M.E. Pollen Analysis (1991), Blackwell Scientific, London 216 pp.
Moreau-Benoit A. Les spores et débris végétaux, Les Schistes et Calcaires éodévoniens de Saint-Cénéré (Massif armoricain, France). Mémoires de la Société géologiques et minéralogique de Bretagne, Rennes (1976) 27-58
Moreau-Benoit A. Les spores et débris végétaux. In: Plusquellec Y. (Ed). Les Schistes et Calcaires de l'Armorique. Mémoires de la Société géologiques et minéralogique de Bretagne, Rennes (1980) 59-78
Moreau-Benoit A., and Dubreuil M. Confirmation et découverte du Dévonien inférieur par la palynoplanctonologie dans les schistes et grès de la terminaison orientale du bassin d'Ancenis (sud-est du Massif armoricain). Géologie de la France 1 (1987) 37-54
Moreau-Benoit A., and Poncet J. Les Schistes et Calcaires éodévoniens de Néhou dans la coupe de Barneville (nord-est du Massif armoricain). Sédimentologie, palynologie, planctonologie, stratigraphie. Bulletin du BRGM 1 2-3 (1982) 81-114
Muñoz J., Felicísimo A.M., Cabezas F., Burgaz A.R., and Martínez I. Wind as a long-distance dispersal vehicle in the Southern Hemisphere. Science 30 (2004) 1144-1147
Powers, R.W., 1968. Lexique stratigraphique international. Volume III, Asie, Fas. 10b1, Arabia Saoudite. Centre Nationale de la Recherche Scientifique, Paris, 177 pp.
Rahmani R.A. Facies and sequences of Devonian Jauf Reservoir, Ghawar Field, Saudi Arabia. 6th Middle East Geosciences Conference, GEO 2004. GeoArabia 9 1 (2004) 118 (Abstract)
Rahmani R.A., Steel R.J., and Al-Duaiji A.A. High-resolution sequence stratigraphy of the Devonian Jauf gas reservoir: a shoreface and estuarine embayment succession, greater Ghawar area, eastern Saudi Arabia. 5th Middle East Geosciencecs Conference, GEO 2002. GeoArabia 7 2 (2002) 288 (Abstract)
Rebske W., Rebske C., Bless M.J.M., Paproth E., and Steemans P. Over enkele fossielen uit de Klerf-Schichten (Onder-Emsium)/Bij Waxweiler (Eifel, BRD). En hun leefmilieu. Grondboor en Hamer 5 (1986) 142-155
Richardson J.B. Chapter 18A. Lower and Middle Palaeozoic records of terrestrial palynomorphs. In: Jansonius J., and McGregor D.C. (Eds). Palynology: Principles and Applications (1996), American Association of Stratigraphic Palynologists Foundation 555-574
Richardson J.B. Taxonomy and classification of some new Early Devonian cryptospores from England. In: Cleal C.J. (Ed). Studies on Early Land Plant Spores from Britain. Special Papers in Palaeontology, Great Britain (1996) 7-40
Richardson J.B., and Edwards D. Sporomorphs and plant megafossils. In: Holland C.H., and Bassett M.G. (Eds). A Global Standard for the Silurian System. Cardiff: National Museum of Wales, Geological Series vol. 9 (1989) 216-226
Richardson J.B., and Ioannides N.S. Silurian palynomorphs from the Tanezzuft and Acacus formations, Tripolitania, North Africa. Micropaleontology 19 (1973) 257-307
Richardson J.B., and Ioannides N.S. Emphanisporites splendens, newname for Emphanisporites pseudoerraticus Richardson and Ioannides, 1973 (preoccupied). Micropaleontology 25 1 (1979) 111
Richardson J.B., and Lister T.R. Upper Silurian and Lower Devonian spore assemblages from the Welsh Borderland and South Wales. Palaeontology 12 (1969) 201-252
Richardson J.B., and McGregor D.C. Silurian and Devonian spore zones of the Old Red Sandstone Continent and adjacent regions. Geological Society of Canada Bulletin 364 (1986) 1-79
Richardson J.B., and Rasul S.M. Palynofacies in a Late Silurian regressive sequence in the Welsh Borderland and Wales. Journal of the Geological Society (London) 147 (1990) 675-686
Richardson J.B., Rasul S.M., and Al-Ameri T. Acritarchs, miospores and correlation of the Ludlovian-Downtonian and Silurian-Devonian boundaries. Review of Palaeobotany and Palynology 34 (1981) 209-224
Richardson J.B., Streel M., Hassan A., and Steemans P. A new spore assemblage to correlate between the Breconian (British Isles) and the Gedinnian (Belgium). Annales de la Société Géologique de Belgique 105 (1982) 135-143
Richardson J.B., Ford J.H., and Parker F. Miospores, correlation and age of some Scottish Lower Old Red Sandstone sediments from the Strathmore region (Fife and Angus). Journal of Micropalaeontolgy 3 2 (1984) 109-124
Richardson J.B., Rodríguez R.M., and Sutherland S.J.E. Palynological zonation of mid-Palaeozoic sequences from the Cantabrian Mountains, NW Spain: implications for inter-regional and interfacies correlation of the Ludford/Prídolí and Silurian/Devonian boundaries, and plant dispersal patterns. Bulletin Natural History Museum London (Geology) 57 2 (2001) 115-162
Roche M., Sabir M., Steemans P., and Vanguestaine M. Palynologie de la région et du sondage de Willerzie. Aarkundige Mededelingen 3 (1986) 149-190
Rodriguez R.M. Mioesporas de la Formación San Pedro/Furada (Silúrico Superior-Devónico Inferior), Cordillera Cantabrica, NO de España. Palinología, Número Extraordinario 1 (1978) 407-433
Rodriguez R.M. Miospores de la Formation San Pedro (Silurien-Devonien) à Corniero (Province de León, Espagne). Revue de Micropaléontologie 20 4 (1978) 216-221
Rodriguez R.M. Nuevas mioesporas de la Formación San Pedro en Geras de Gordon, Cordillera Cantabrica (Provincia de León, Noroeste de España). Breviora Geologica Asturica 22 1-2 (1978) 9-16
Rodriguez R.M. Palinologia de las Formaciones del Silúrico superior-Devonico inferior de la Cordillera Cantabrica (1983), Servicio de Publicaciones, Universidad de Leon 1-231
Rubinstein C.V., and Steemans P. Miospore assemblages from the Silurian-Devonian boundary, in borehole A1-61, Ghadamis Basin, Libya. In: Steemans P., Servais T., and Streel M. (Eds). Paleozoic Palynology: A Special Issue in Honour of Dr. Stanislas Loboziak. Review of Palaeobotany and Palynology, Amsterdam (2002) 397-421
Rubinstein C.V., Melo J.H.G., and Steemans P. Lochkovian (earliest Devonian) miospores from the Solimões Basin, northern Brazil. Review of Palaeobotany and Palynology 133 (2005) 91-113
Rutty P.M., Rahmani R.A., Steel R.J., Hulver M., Johnson R.S., and Demaree R.G. Saudi Arabian Silurian and Devonian sequence stratigraphy, paleogeography and play concepts. 5th Middle East Geosciences Conference, GEO 2002. GeoArabia 7 2 (2002) 291 (Abstract)
Sharland P.R., Archer D.M., Casey R.B., Davies S.H., Hall A.P., Heward A.D., Horbury A.D., and Simmons M.D. Arabian plate sequence stratigraphy. GeoArabia, Special Publication 2 Gulf PetroLink, Manama, Bahrain (2001) 371 pp.
Steemans P. Etude stratigraphique des spores dans les couches de transition "Gedinnien-Siegenien" à Nonceveux et à Spa (Belgique). Annales de la Société Géologique de Belgique 104 (1981) 41-59
Steemans P. Gedinnian and Siegenian spores stratigraphy in Belgium. In: Ziegler W., and Werner R. (Eds). On Devonian Stratigraphy and Paleontology of the Ardenno-Rhenish Mountains and Related Devonian Matters. Courier Forschungsinstitut Senckenberg vol. 55 (1982) 165-180
Steemans, P., 1982b. L'âge du poudingue de Fépin (base du Gedinnien) à Lahonry (Belgique). Bulletin de la Société belge de Géologie (1981)90(4): 331-340.
Steemans P. Révision palynostratigraphique de l'âge des quartzites blancs de la coupe du Wayai (Km 7). Massif de Theux-Belgique. Annales de la Société Géologique de Belgique 110 (1988) 363-369
Steemans P. Paléogéographie du bassin ardennais au cours de l'Eodévonien. Annales de la Société Géologique de Belgique 112 1 (1989) 103-119
Steemans P. Palynostratigraphie de l'Eodévonien dans l'ouest de l'Europe. Mémoires Explicatifs pour les Cartes Géologiques & Minéralogiques de la Belgique, 27 (1989), Service Géologique de Belgique, Bruxelles 453 pp.
Steemans P. Silurian and Lower Emsian spores in Saudi Arabia. Review of Palaeobotany and Palynology 89 (1995) 91-104
Steemans P. Cryptospores and spores from the Ordovician to the Llandovery. A review. Acta Universitatis Carolinae - Geologica 43 1/2 (1999) 271-273
Steemans P. Paléodiversification des spores et des cryptospores de l'Ordovicien au Dévonien inférieur. Géobios 32 2 (1999) 341-352
Steemans P., and Gerrienne P. La micro- et macroflore du Gedinnien de la Gileppe, Synclinorium de la Vesdre, Belgique. Annales de la Société Géologique de Belgique 107 (1984) 51-71
Steemans P., and Lakova I. The Moesian Terrane during the Lochkovian - a new palaeogeographic and phytogeographic hypothesis based on miospore assemblages. Palaeogeography, Palaeoclimatology, Palaeoecology 208 3-4 (2004) 225-233
Steemans P., and Wellman C.H. Miospores and the emergence of land plants. In: Webby B., Paris F., Droser M.L., and Percival I.G. (Eds). The Great Ordovician Biodiversification Event. Critical Moments and Perspectives in Earth History and Paleobiology (2004), Columbia University Press, New York 361-366
Steemans P., Le Hérissé A., and Bozdogan N. Ordovician and Silurian cryptospores and miospores from Southeastern Turkey. Review of Palaeobotany and Palynology 93 (1996) 35-76
Steemans P., Debbaut V., and Faber A. Preliminary survey of the palynological content of the Lower Devonian in the Oesling, Luxembourg. Bulletin de la Société des Naturalistes Luxembourgeois 100 (2000) 171-186
Steemans P., Al-Ruwaili M., Filatoff J., and Streel M. A miospore assemblage at the Siluro-Devonian boundary from the Tawil Formation, Qatif Field, Saudi Arabia. In: McLean D. (Ed). First Meeting of the CIMP, Spores and Pollen Subcommission (2001), National University of Ireland, Cork 43 (2-7 September)
Steemans, P., Rubinstein, C., Melo, J.H.G., in press. Miospore biostratigraphy of the Siluro-Devonian boundary, Urubu area, Western Amazonas Basin, Northern Brazil. Geobios.
Streel M. Associations de spores du Dévonien inférieur belge et leur signification stratigraphique. Annales de la Société Géologique de Belgique 90 Bull. 1-3 (1967) 11-53
Streel M., Fairon-Demaret M., Otazo-Bozo N., and Steemans P. Etude stratigraphique des spores du Dévonien inférieur au bord sud du Synclinorium de Dinant (Belgique) et leurs applications. Annales de la Société Géologique de Belgique 104 (1981) 173-171
Streel M., Higgs K., Loboziak S., Riegel W., and Steemans P. Spore stratigraphy and correlation with faunas and floras in the type marine Devonian of the Ardenno-Rhenish regions. Review of Palaeobotany and Palynology 50 (1987) 211-229
Streel M., Steemans P., and Loboziak S. Devonian miospore stratigraphy in Libya and surrounding areas compared with related faunal and floral data. International symposium circum-Mediterranean palynology, Zeist (1988) April 19-23
Streel M., Fairon-Demaret M., Gerrienne P., Loboziak S., and Steemans P. Lower and Middle Devonian miospore-based stratigraphy in Libya and its relation to the megafloras and faunas. Review of Palaeobotany and Palynology 66 3/4 (1990) 229-242
Streel M., Fairon-Demaret M., and Loboziak S. Givetian-Frasnian phytogeography of Euramerica and western Gondwana based on miospore distribution. In: McKerrow W.S., and Scotese C.R. (Eds). Palaeozoic Palaeogeograpphy and Biogeography (1990), Geological Society 291-296
Strother P.K. A classification schema for the cryptospores. Palynology 15 (1991) 219-236
Stump T.E., and Van der Eem J.G. In: Al-Husseini M.I. (Ed). Overview of the stratigraphy, depositional environments and periods of deformation of the Wajid outcrop belt, southwestern Saudi Arabia. The Middle East Petroleum Geosciences, Geo '94 vol. 2 (1994), Gulf PetroLink, Manama, Bahrain 867-876
Stump T.E., and Van der Eem J.G. The stratigraphy, depositional environments and periods of deformation of the Wajid outcrop belt, southwestern Saudi Arabia. Journal of African Earth Sciences 21 3 (1995) 421-441
Stump T.E., Al-Hajri S., and Van der Eem J.G.L.A. Geology and biostratigraphy of the Late Precambrian through Palaeozoic sediments of Saudi Arabia. Review of Palaeobotany and Palynology 89 1/2 (1995) 5-17
Tauber H. Differential pollen dispersion and the interterpretation of pollen diagrams. Danmarks Geologiske Undersølgelse 89 2 (1965) 1-69
Taugourdeau-Lantz P., and de Jekhowsky B. Répartition et description des chitinozoaires siluro-dévoniens de quelques sondages de la C.R.E.P.S., de la C.F.P.A. et de la S.N. Repal au Sahara. Revue de l'Institut Français du Pétrole et Annales des combustibles liquides 15 9 (1960) 1199-1260
Traverse A. Paleopalynology (1988), Unwin Hyman, Boston 600 pp.
Tryon R. Development and evolution of fern floras of oceanic islands. Biotropica 2 (1970) 76-84
Tryon R. Endemic areas and geographic speciation in tropical American ferns. Biotropica 4 3 (1972) 121-131
Tuomisto H., Ruokolainen K., and Yli-Halla M. Dispersal, environment, and floristic variation of Western Amazonian forests. Science 299 (2003) 241-244
Turnau E. Lower to Middle Devonian spores from the vicinity of Pionki (Central Poland). Review of Palaeobotany and Palynology 46 (1986) 311-354
Turnau E. Miospore stratigraphy of Middle Devonian deposits from Western Pomerania. Review of Palaeobotany and Palynology 93 1-4 (1996) 107-125
Turnau E., and Jakubowska L. Early Devonian miospores and age of the Zwolen Formation (Old Red Sandstone facies) from Ciepielów IG-1 borehole. Annales Societatis Geologorum Poloniae 59 (1989) 391-416
Turnau E., and Matyja H. Timing of the onset of Devonian sedimentation in Northwestern Poland: palynological evidence. Annales Societatis Geologorum Poloniae 71 (2001) 67-74
Turnau E., and Racki G. Givetian palynostratigraphy and palynofacies: new data from the Bodzentyn Syncline (Holy Cross Mountains, central Poland). Review of Palaeobotany and Palynology 106 3-4 (1999) 237-271
Van Zanten B.O. Experimental studies on trans-oceanic long-range dispersal of moss spores in the southern hemisphere. Journal of the Hattori Botanical Laboratory 44 (1978) 445-482
Vigran J.O. Spores from Devonian deposits, Mimerdalen, Spitsbergen. Norsk Polarinstitutt 132 (1964) 1-32
Van Zanten B.O., and Gradstein S.R. Experimental dispersal geography of neotropical liverworts. Beiheft zue Nova Hedwigia 90 (1988) 41-94
Van Zanten B.O., and Pócs T. Distribution and dispersal of bryophytes. Advances in Bryology 1 (1981) 479-562
Verniers J., Nestor V., Paris F., Dufka P., Sutherland S., and Van Grootel G. A global Chitinozoa biozonation for the Silurian. Geological Magazine 132 6 (1995) 651-666
Wallace, C.A., Dini, S.M. and Al-Farasani, A.A., 1996. Explanatory notes to the geological map of the Ash Shuwahitiya Quadrangle, Kingdom of Saudi Arabia. Geoscience Map GM-125C, scale 1:250,000, sheet 30D. Deputy Ministry for Mineral Resources, Ministry of Petroleum and Mineral Resources, Kingdom of Saudi Arabia, 27 pp.
Wallace, C.A., Dini, S.M., Al-Farasani, A.A., 1997. Explanatory notes to the geological map of the Al Jawf Quadrangle, Kingdom of Saudi Arabia. Geoscience Map GM-128C, scale 1:250,000, sheet 29D. Deputy Ministry for Mineral Resources, Ministry of Petroleum and Mineral Resources, Kingdom of Saudi Arabia, 32 pp.
Wellman C.H. A Lower Devonian sporomorph assemblage from the Midland Valley of Scotland. Transactions of the Royal Society of Edinburgh. Earth Sciences 84 (1993) 117-136
Wellman C.H., and Gray J. The microfossil record of early land plants. Philosophical Transactions of the Royal Society of London - Series B. Biological Sciences 355 1398 (2000) 717-731 (discussion 731-2)
Wellman C.H., and Richardson J.B. Terrestrial plant microfossils from Silurian inliers of the Midland Valley of Scotland. Palaeontology 36 1 (1993) 155-193
Wellman C.H., and Richardson J.B. Sporomorph assemblages from the 'Lower Old Red Sandstone' of Lorne Scotland. In: Cleal C.J. (Ed). Studies on Early Land Plant Spores From Britain. Special Papers in Palaeontology, Great Britain (1996) 41-101
Wellman C.H., Habgood K., Jenkins G., and Richardson J.B. A new assemblage (microfossil and megafossil) from the Lower Old Red Sandstone of the Anglo-Welsh Basin: its implications for the palaeoecology of early terrestrial ecosystems. Review of Palaeobotany and Palynology 109 (2000) 161-196
Wender L.E., Bryant J.W., Dickens M.F., Neville A.S., and Al-Moqbel M.A. Paleozoic (pre-Khuff) hydrocarbon geology of the Ghawar area, eastern Saudi Arabia. GeoArabia 3 2 (1998) 273-302
Wild M., and Gagnon D. Does lack of suitable habitat explain the patchy distributions of rare calcicole fern species?. Ecography 28 2 (2005) 191-196
Wood G.D., and Benson D.G. The North American occurrence of the algal coenobium Plaesiodictyon: paleogeographic, paleoecologic, and biostratigraphic importance in the Triassis. Palynology 24 (2000) 9-20